Cleretum bellidiforme leaves

The small papillae on Cleretum bellidiforme leaves vary in size on different leaves. On some leaves their distribution may be random, on others they are arranged in longitudinal rows as in the photo.

Some sand has accumulated on the leaves in picture. It is never too far from any plant parts, given where they grow and how low they become (Vlok and Schutte-Vlok, 2015; Manning, 2007, iNaturalist).